Principal Engineer, AI Security

About this role

Your Impact at LILA

As a Principal Security Engineer focused on AI Security, you will define and drive the technical strategy for securing how AI is used across Lila's enterprise. You will operate as a senior individual contributor, partnering with IT and business teams to ensure safe and compliant adoption of AI tools and platforms.

While Lila builds AI-powered systems, this role is primarily focused on securing the use of third-party and internally deployed AI tools across the enterprise — ensuring sensitive data, intellectual property, and scientific workflows are protected as AI becomes deeply embedded in how work gets done.

What You'll Be Building

• Enterprise AI Security Strategy — Define and implement security controls and guardrails for the use of AI tools (e.g., LLM APIs, SaaS AI platforms, and internal AI services) across the organization.

• AI Gateway & Agentic Gateway Security — Design and implement AI gateway controls to manage and monitor access to external and internal AI systems. Secure agentic workflows by enforcing identity, authorization, tool-use constraints, and policy controls for autonomous or semi-autonomous agents.

• AI Red Teaming & Adversarial Testing — Conduct red teaming and adversarial testing focused on enterprise AI usage, including prompt injection, data exfiltration, jailbreaks, and abuse of connected tools and plugins.

• Data Protection for AI Usage — Develop and enforce controls to prevent sensitive data leakage through AI systems, including input/output filtering, data classification, tokenization, and secure handling of prompts, embeddings, and outputs.

• Multi-Layer AI Security (Network, Endpoint, Data) — Integrate AI security into existing enterprise security layers: network visibility and control over AI service access, API traffic inspection, and zero trust enforcement; endpoint security for developer machines, research environments, browsers, and plugins; data layer controls ensuring proper handling of sensitive data when interacting with AI systems.

• AI Threat Modeling (Enterprise Context) — Develop threat models focused on enterprise AI usage, including risks such as data leakage, prompt injection, model misuse, supply chain risks from AI vendors, and unauthorized agent actions.

• Vendor & Platform Security — Assess and guide secure adoption of third-party AI vendors and platforms, including evaluating data handling practices, model behavior, and integration risks.

• Incident Response for AI Usage — Define and support response approaches for AI-related incidents, such as sensitive data exposure, policy violations, or misuse of AI tools.

• Cross-Functional Technical Leadership — Partner with Legal, Compliance, IT, and Engineering to align AI usage with regulatory requirements, data governance policies, and responsible AI practices.

• Security Enablement — Contribute to internal guidance and education on safe AI usage, including secure prompting, data handling, and appropriate use of AI tools.

• Security Tooling & Implementation — Evaluate and implement tooling for AI security, including AI gateways, DLP integrations, monitoring solutions, and policy enforcement mechanisms.

What You’ll Need to Succeed

• 8+ years of experience in information security, with strong expertise in enterprise, cloud, or application security.

• Hands-on experience designing and implementing security controls in enterprise environments.

• Familiarity with AI/ML systems and how modern AI tools (LLMs, copilots, APIs) are used in practice.

• Experience with cloud platforms (AWS/GCP), SaaS security, and zero trust architectures.

• Experience with data protection technologies (e.g., DLP, data classification, access controls).

• Practical experience with threat modeling, red teaming, or adversarial testing.

• Strong communication and influence skills across technical and non-technical stakeholders.

Bonus Points For

• Experience securing enterprise use of LLMs, copilots, or generative AI platforms.

• Familiarity with AI gateways, prompt filtering, or model interaction controls.

• Experience evaluating or securing third-party AI vendors and APIs.

• Background in regulated environments (biotech, healthcare, defense, or government).

• Experience with browser security, endpoint controls, or SaaS security platforms.

• Knowledge of privacy-enhancing technologies or confidential computing.

• Contributions to AI/ML security research or community.

Compensation

We offer competitive base compensation with bonus potential and generous early-stage equity. Your final offer will reflect your background, expertise, and expected impact.

U.S.…